diff --git a/apex/apex.go b/apex/apex.go
index 46aaa8b5f..c31ed765e 100644
--- a/apex/apex.go
+++ b/apex/apex.go
@@ -2311,7 +2311,10 @@ func (a *apexBundle) GenerateAndroidBuildActions(ctx android.ModuleContext) {
 	// Build rules are generated by the dexpreopt singleton, and here we access build artifacts
 	// via the global boot image config.
 	if a.artApex {
-		for arch, files := range java.DexpreoptedArtApexJars(ctx) {
+		artAndOatFiles, vdexFiles := java.DexpreoptedArtApexJars(ctx)
+
+		// Copy *.art and *.oat files to arch-specific subdirectories.
+		for arch, files := range artAndOatFiles {
 			dirInApex := filepath.Join("javalib", arch.String())
 			for _, f := range files {
 				localModule := "javalib_" + arch.String() + "_" + filepath.Base(f.String())
@@ -2319,6 +2322,18 @@ func (a *apexBundle) GenerateAndroidBuildActions(ctx android.ModuleContext) {
 				filesInfo = append(filesInfo, af)
 			}
 		}
+
+		// Copy *.vdex files to a common subdirectory.
+		for _, file := range vdexFiles {
+			dirInApex := "javalib"
+			localModule := "javalib_" + filepath.Base(file.String())
+			af := newApexFile(ctx, file, localModule, dirInApex, etc, nil)
+			// Add a symlink to the *.vdex file for each arch-specific subdirectory.
+			for arch := range artAndOatFiles {
+				af.symlinks = append(af.symlinks, filepath.Join(arch.String(), filepath.Base(file.String())))
+			}
+			filesInfo = append(filesInfo, af)
+		}
 	}
 
 	if a.private_key_file == nil {
diff --git a/apex/builder.go b/apex/builder.go
index 67bc20603..ea1215e1e 100644
--- a/apex/builder.go
+++ b/apex/builder.go
@@ -333,6 +333,7 @@ func (a *apexBundle) buildUnflattenedApex(ctx android.ModuleContext) {
 	for _, fi := range a.filesInfo {
 		destPath := android.PathForModuleOut(ctx, "image"+suffix, fi.Path()).String()
 		copyCommands = append(copyCommands, "mkdir -p "+filepath.Dir(destPath))
+
 		if a.linkToSystemLib && fi.transitiveDep && fi.AvailableToPlatform() {
 			// TODO(jiyong): pathOnDevice should come from fi.module, not being calculated here
 			pathOnDevice := filepath.Join("/system", fi.Path())
@@ -341,10 +342,16 @@ func (a *apexBundle) buildUnflattenedApex(ctx android.ModuleContext) {
 			copyCommands = append(copyCommands, "cp -f "+fi.builtFile.String()+" "+destPath)
 			implicitInputs = append(implicitInputs, fi.builtFile)
 		}
+
 		// create additional symlinks pointing the file inside the APEX
 		for _, symlinkPath := range fi.SymlinkPaths() {
 			symlinkDest := android.PathForModuleOut(ctx, "image"+suffix, symlinkPath).String()
-			copyCommands = append(copyCommands, "ln -sfn "+filepath.Base(destPath)+" "+symlinkDest)
+			symlinkTarget, err := filepath.Rel(filepath.Dir(symlinkDest), destPath)
+			if err != nil {
+				panic("Cannot compute relative path from " + destPath + " to " + filepath.Dir(symlinkDest))
+			}
+			copyCommands = append(copyCommands, "mkdir -p "+filepath.Dir(symlinkDest))
+			copyCommands = append(copyCommands, "ln -sfn "+symlinkTarget+" "+symlinkDest)
 		}
 	}
 
@@ -407,6 +414,7 @@ func (a *apexBundle) buildUnflattenedApex(ctx android.ModuleContext) {
 				}
 			} else {
 				readOnlyPaths = append(readOnlyPaths, pathInApex)
+				readOnlyPaths = append(readOnlyPaths, f.SymlinkPaths()...)
 			}
 			dir := f.installDir
 			for !android.InList(dir, executablePaths) && dir != "" {
diff --git a/java/dexpreopt_bootjars.go b/java/dexpreopt_bootjars.go
index a3b264ed8..0751c0862 100644
--- a/java/dexpreopt_bootjars.go
+++ b/java/dexpreopt_bootjars.go
@@ -186,19 +186,31 @@ type dexpreoptBootJars struct {
 }
 
 // Accessor function for the apex package. Returns nil if dexpreopt is disabled.
-func DexpreoptedArtApexJars(ctx android.BuilderContext) map[android.ArchType]android.OutputPaths {
+func DexpreoptedArtApexJars(ctx android.BuilderContext) (map[android.ArchType]android.OutputPaths, android.OutputPaths) {
 	if skipDexpreoptBootJars(ctx) {
-		return nil
+		return nil, nil
 	}
-	// Include dexpreopt files for the primary boot image.
-	files := map[android.ArchType]android.OutputPaths{}
-	for _, variant := range artBootImageConfig(ctx).variants {
+
+	image := artBootImageConfig(ctx)
+
+	// Target-independent boot image files (*.vdex).
+	anyTarget := image.variants[0].target
+	vdexDir := image.dir.Join(ctx, anyTarget.Os.String(), image.installSubdir, anyTarget.Arch.ArchType.String())
+	vdexFiles := image.moduleFiles(ctx, vdexDir, ".vdex")
+
+	// Target-specific boot image files (*.oat, *.art).
+	artAndOatFiles := map[android.ArchType]android.OutputPaths{}
+	for _, variant := range image.variants {
 		// We also generate boot images for host (for testing), but we don't need those in the apex.
-		if variant.target.Os == android.Android {
-			files[variant.target.Arch.ArchType] = variant.imagesDeps
+		os := variant.target.Os
+		if os == android.Android {
+			arch := variant.target.Arch.ArchType
+			archDir := image.dir.Join(ctx, os.String(), image.installSubdir, arch.String())
+			artAndOatFiles[arch] = image.moduleFiles(ctx, archDir, ".art", ".oat")
 		}
 	}
-	return files
+
+	return artAndOatFiles, vdexFiles
 }
 
 // dexpreoptBoot singleton rules